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What is the target audience of these directories?

These directories, being general, English web directories, target a global, English speaking audience. Since you can choose the way directories link to your site, you could use titles to target specific keywords or phrases with which you would expect to be searched. This will increase the probability of your site ranking high for those targeted terms or phrases used in your titles, which in turn could lead to you reaching out to your target audience (who conduct the searches).

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